Columns from left to right (omitting blanks) are:
Tiranige (transcription)
finder (English finder list, i.e. usually one-word glosses)
recherche (finder list in French)
English (full English gloss)
français (full French gloss)
code (for certain lexical categories: fauna, flora, body, kinship)
order/family (for flora-fauna terms)
species (for flora-fauna terms)
synonymy (alternative or former species binomials)